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Ginger "Gr" : (this may not be a distinct gene). This gene may be Columbian "Co" or closely related Dark brown "Db".
Columbian "Co" Incompletely dominant. Confines black to hackle and tail in both sexes (called Columbian restriction). Thought to cause a gradient in color from head to tail. Modifies Wheaten to Buff Columbian. Has no effect on extended black, E.
Dark brown "Db" Incompletely dominant. Changes black down of E, ER to reddish-brown. Adults males exhibit a Columbian-type pattern of black, modifies red to orange-tan. Db is a better restrictor of black in males than females.